Репродуктивное поведение сивуча
Диссертация
Впервые при помощи нового метода, сканирующего фотои видеорегистрации событий и мест положения животных на лежбище, удалось детально количественно описать и показать, как организованы репродуктивные процессы на лежбище, как совместно действуют система репродуктивных связей между самцами и самками и система связей между матерью и детенышем, а также как самка использует репродуктивные стратегии… Читать ещё >
Содержание
- 1. 1. Цель работы
- 1. 2. Задачи
- 1. 3. Научная новизна
- 1. 4. Теоретическое и практическое значение результатов работы
- 2. 1. Биология и годовой цикл жизни
- 2. 2. Полигиния у сивучей
- 2. 3. Территориальность сивучей
- 2. 3. 1. Самцы
- 2. 3. 2. Самки
- 2. 4. Репродуктивный успех самцов и самок
- 2. 5. Территориальность вне репродуктивных скоплений
- 2. 6. Режим использования репродуктивного лежбища самками
- 3. 1. Идентификация и регистрация меченых и приметных животных
- 3. 1. 1. Регистрация основных событий для меченых и приметных сивучей
- 3. 2. Методы сбора пространственных данных
- 3. 2. 1. Модельные участки
- 3. 2. 2. Обработка изображений
- 3. 2. 3. Преобразование координат событий
- 3. 2. 4. Оценка ошибки метода преобразования координат событий
- 3. 2. 5. Объем материала
- 3. 2. 6. Обработка пространственных данных
- 3. 3. Методы визуальных наблюдений
- 3. 3. 1. Продолжительность использования репродуктивного пространства лежбища
- 3. 3. 2. Исследование режима присутствия самок, регистрация приходов-уходов
- 3. 3. 3. Факторы, определяющие территориальное поведение самцов
- 4. 1. Временные характеристики формирования и распада репродуктивных скоплений
- 4. 2. Временные характеристики периодов родов и спариваний
- 4. 3. Режим присутствия самок на лежбище
- 4. 3. 1. Суточная динамика вероятности присутствия
- 4. 3. 2. Суточные ритмы уходов и возвращений самок на лежбище
- 4. 3. 3. Продолжительность периодов присутствия и отсутствия самок на лежбище
- 4. 4. Использование пространства репродуктивного участка самками сивуча
- 4. 5. Использование пространства лежбища территориальными самцами
- 4. 6. Число репродуктивных сезонов, в которых секач удерживает территорию на лежбище
- 4. 7. Оценка уровня дисперсии самцов и самок, рожденных на Скале Долгая
- 4. 8. Спаривания и роды в пространстве лежбища
- 4. 8. 1. Сезонная динамика репродуктивных процессов в пространстве
- 4. 8. 2. Межгодовые тенденции протекания репродуктивных процессов в пространстве
- 4. 9. Анормальное копуляторное поведение секачей
- 4. 10. Копуляторный успех самцов в зависимости от положения в пространстве лежбища
- 4. 11. Территориальное поведение самцов на нерепродуктивном участке лежбища
- 4. 11. 1. Демография Главного Холостякового участка
- 4. 11. 2. Характер залегания животных различных половозрастных групп
- 4. 11. 3. Бюджет времени самцов различного возраста
- 4. 11. 4. Сравнение территориальной активности самцов до и после прихода самок
- 5. 1. Временные характеристики использования пространства лежбища сивучами
- 5. 2. Режим присутствия самок сивуча на лежбище
- 5. 2. 1. Различия в режимах пребывания самок сивуча на берегу и уходов на фуражирование
- 5. 2. 2. Чем можно объяснить различия
- 5. 3. Использование пространства лежбища сивучами
- 5. 3. 1. Самки и самцы в пространстве лежбища
- 5. 4. Спаривания и роды в пространстве лежбища
- 5. 5. Репродуктивные стратегии самцов
- 5. 5. 1. Структура репродуктивных связей
- 5. 6. Территориальное поведение и его проявление у самцов разного возраста на нерепродуктивных участках лежбища
Список литературы
- Андрияшев, А., 1979. О некоторых вопросах вертикальной зональности донной морской фауны. В кн.: Биологические ресурсы Мирового океана. М.: Наука. С. 117−137.1. Дисс. канд. биол. наук
- Белкин, А.Н., 1966. Летнее распределение, запасы, перспективы промысла и некоторые черты биологии сивуча, обитающего на Курильских островах. // Изв. ТИНРО. Т. 58. С. 69−95.
- Владимиров В.А., Лыскин Н. Н. 1984. Новые данные по размножению и структуре популяций северных котиков (Callorhinus иг sinus L.) // Зоол. журн. Т. 63. Вып. 12. С. 1883−1890.
- Капитонов, Ю., Палагин, Ю., Шалыгин, А., 1988. Восстановление плотности распределения случайных полей и процессов по экспериментальным данным с помощью ядерных оценок //Автометрия. Т. 6. 49 с.
- Крушинская Н.Л., Лисицына. Т.Ю., 1983. Поведение морских млекопитающих. М.: Наука. 336 с.
- Кузин, А., 1996. Численность, перемещения и основные черты биологии сивуча о. Тюленъего//Изв. ТИНРО. Т. 121. С. 130−142.
- Кузин, А.Е., 2005. О продолжительности пребывания на берегу и в море сивучей о. Тюленьего (Охотское море). //Изв. ТИНРО. Т. 140. С. 108−117.
- Лисицына, Т., 1979. Звуковая сигнализация в поведении самцов сивучей (Eumetopias jubatus) на лежбищах // Зоол. ж. Т. 58. № 10. С. 1552−1557.
- Лисицына, Т., 1976. Территориальное поведение сивучей Eumetopias jubatus (Otariidae) //Зоол. ж. Т. 55. № 3. Р. 408−420.
- Лисицына, Т., 1975. Лежбища сивуча на острове Ионы //Тез. докл. VIВсес. совещ., Киев. С. 179−181.
- Лобачева, Н.В., 1998. Отношения мать-детеныш у сивуча на Юго-Восточном лежбище о. Медный. Дипломная работа: МГУ им. М. В. Ломоносова, биологический факультет. 56 с.
- Мамаев, Е.Г., 1997. Бюджет времени и активность секачей сивуча Eumetopias jubatus (Pinnipedia, Otariidae) на репродуктивном лежбище // Зоол. ж. Т. 76 № 4. Р. 494−500.
- Мамаев, Е., 1999. Территориальное поведение секачей сивуча Eumetopias jubatus в репродуктивный период. Дисс. канд. биол. наук. М: МГУ им. М. В. Ломоносова, биологический факультет. 177с. woe поведение сиеуча
- Мараков, С.В., 1966. Морской лев—сивуч //Природа. Т. 8. С. 61−73.
- Мужчинкин, В., 1964. Распределение сивучей на Юго-Восточном котиковом лежбище острова Медного. //Изв. ТИНРО. Т. 54. С. 179−186.
- Перлов, А., 1983. Отношение ко льдам ушастых тюленей //Биологические проблемы севера. Тезисы X Всесоюзного симпозиума (Магадан). С. 129−130.
- Перлов, А., 1980. Возрастной состав и соотношение полову сивучей, Eumetopias jubatus (Otariidae) на лежбищах Курильских островов // Зоол. ж. Т. 59. С. 1545−1553.
- Перлов, А., 1971. Сроки наступления половой зрелости у сивучей. // Тр. ВНИРО / Изв. ТИНРО. Т. 82 / 80. С. 174−189.
- Попов Л. А. 1982. К биологии сивуча Берингова моря // Изучение, охрана и рациональное использование морских млекопитающих: Тез. Докл. VIII Всесоюзного совещания — Астрахань. С. 294−295.
- Челноков, Ф.Г., 1985. О протяженности и числе лежбищ на Командорских островах в прошлое и настоящее время//Вопросы географии Камчатки. Т. 9. С. 106−109.
- Чугунков, Д.И., 1968. Сивучи и котиковое хозяйство Командорских островов //Изв. ТИНРО. Т. 68. С. 159−171.
- Allee, W.C., 1951. Co-operation among animals, with human implications. N.Y.: Henry Shuman. 233 pp.
- Allee, W., 1952. Dominance and hierarchy in societies of vertebrates // Colloques internationaux du CNRS. V. 34. P. 157−181.
- Allen, J., 1880. History of North American pinnipeds: a monograph of the walruses, sealions, sea-bears and seals of North America. GPO. (ISBN: 12)
- Altmann.J., 1974. Observational study of behavior: sampling methods // Behaviour. V. 49(3). P. 227−267. JSTOR.
- Altukhov, A. & Burkanov, V., 2008. Steller Sea Lion Brand Resight Database Using MS Access. // Alaska Marine Science Symposium, Anchorage, Alaska, USA, 20−23 January, 2008. P. 35.
- Altukhov, A. V. & Burkanov, V.N., 2008. Abnormal mating behavior of Steller sea lion bulls// 22th Annual Conference of European Cetacean Society. Marine Mammals in Time: Past, Present and Future. 10−12 March, Egmond Aan Zee, The Netherlands. P. 98.
- Altukhov, A. & Burkanov, V., 2009. Survival and emigration rate estimations of Steller sea lions (Eumetopias jubatus) on the Dolgaya Rock rookery (Kuril Islands, Russia) // 18thflucc. K3H0. 6uon, Haya
- Biennial Conference on the Biology of Marine Mammals, Quebec City, Quebec, 12−16 October 2009. P. 13.
- Ambs, S., Boness, D., Bowen, W., Perry, E. & Fleischer, R., 1999. Proximate factors associated with high levels of extraconsortfertilization inpolygynous grey seals //Animal Behaviour. V. 58(3). P. 527−535. Elsevier.
- Amos, W., Wilmer, J.W. & Kokko, H., 2001. Do female grey seals select genetically diverse mates? (Halichoerus grypus) // Animal Behaviour. V. 62(1). P. 157−164.
- Amos, B., Twiss, S., Pomeroy, P. & Anderson, S., 1995. Evidence for mate fidelity in the gray seal. (Halichoerus grypus) //Science. V. 268(5219). P. 1897−1899.
- Andrew, T. & Robertson, R., 1984. Aggressive and territorial behaviour in female red-winged blackbirds // Canadian journal of zoology. V. 62(2). P. 148−153. NRC Research Press.
- Appleby, M.C., 1982. The consequences and causes of high social rank in red deer stags. // Behaviour. V. 80. P. 259−273.
- Arak, A. 1984. Sneaky breeders. In: Producers and scroungers: strategies of exploitation and parasitism. Taylor & Francis, 1984. P. 154−194.
- Arnould, J.P.Y. & Duck, C.D., 1997. The cost and benefits of territorial tenure, and factors affecting mating success in male Antarctic fur seals. (Arctocephalus gazella) // Journal of Zoology. V. 241(4). P. 649−664.
- Baker, J.D., Antonelis, G.A., Fowler, C.W. & York, A.E., 1995. Natal site fidelity in northern fur seals, Callorhinus ursinus. //Animal Behaviour. V. 50(1). P. 237−247.
- Baker, A.R.T.R.L.V.B.C.W.M.R.G.T.D.G.C.J.K.W. & Bickham, J.W., 2005. Variation of mitochondrial control region sequences of Steller sea lions: The three-stock hypothesis. // Journal of Mammalogy. V. 86(6). P. 1075−1084.
- Barnett, S.A., 1958. An analysis of social behaviour in wild rats. //Proceedings of the Zoological Society of London. V. 130. P. 107−152.
- Bartholomew, G., 1953. Behavioral factors affecting social structure in the Alaska fur seal// Trans. N. Am. Wildl. Corf. P. 481−501.
- Bartholomew, G., 1970. A model for the evolution ofpinniped polygyny// Evolution. V. 24. P. 546−559.
- Beehler, B. & Foster, M., 1988. Hotshots, hotspots, and female preference in the organization of lek mating systems // American Naturalist. V. 131. P. 203−219. JSTOR.
- Bermond, B., Mos, J., Meelis, W" Van Der Poel, A.M. & Kruk, M.R., 1982. Aggression induced by stimulation of the hypothalamus: effects of androgens. //Pharmacology, Biochemistry and Behavior. V. 16. P. 41−45.
- Birch, C., 1999. A new generalized logistic sigmoid growth equation compared with the Richards growth equation// Annals of Botany. V. 83(6). P. 713. Annals Botany Co.
- Boness, D.J., 1991. Determinants of mating systems in the Otariidae (Pinnipedia). In: Behaviour of Pinnipeds. RenoufD. (ed). London: Chapman and Hall Ltd. P. 410. (ISBN: 0412−30 540−2)
- Boness, D.J. & James, H., 1979. Reproductive behaviour of the grey seal (Halichoerus grypus) on Sable Island, Nova Scotia. // Journal of Zoology. V. 188(4). P. 477−5
- Boyd, I., Arnould, J., Barton, T. & Croxall, J., 1994. Foraging behaviour of Antarctic fur seals during periods of contrasting prey abundance. //Journal of Animal Ecology. V. 63. P. 703−713.
- Boyd, I.L., 1996. Individual variation in the duration of pregnancy and birth date in Antarctic fur seals: The role of environment, age, and sex offetus. (Arctocephalus gazella) // Journal of Mammalogy. V. 77(1). P. 124−133.
- Boyd, I.L. & Duck, C.D., 1991. Mass changes and metabolism in territorial male Antarctic fur seals (Arctocephalus gazella). //Physiological Zoology. V. 64. P. 375−392.
- Bradbury, J. W. 1981. The evolution ofleks. // Natural selection and social behavior. Chiron Press, New York. P. 138−169.1. Дисс. канд. биоп. наук
- Bradbury, J. W. & Gibson, R.M., 1983. Leks and mate choice. In: Mate choice. Bateson, P.P.G. (ed). Cambridge University Press. P. 109−138. (ISBN: 0−521−27 207−6)
- Brandon E.A., 1997. Steller Sea Lion Research Peer Review: Behavior// Rookery Studies Workshop, Seattle, Washington, December 5−7. P. 16—17.
- Brandon, E. A. 2000. Maternal investment in Steller sea lions in Alaska. Ph.D. thesis, Texas A&M University, Galveston, TX. 137pp.
- Bronson, F.H., 1981. Mammalian reproduction: an ecological perspective // Biology of Reproduction. V. 32. P. 1−26.
- Burkanov, V. 2009, National Marine Mammal Laboratory (NMML), Alaska Ecosystem Program. «Russian Steller Sea Lion Research Update"//AFSC Quarterly report, 2009. P.6−11.
- Burt, W., 1943. Territoriality and home range concepts as applied to mammals//Journal of Mammalogy. V. 24(3). P. 346−352. JSTOR.
- Calenge, C., 2006. The package «adehabitat» for the R software: A tool for the analysis of space and habitat use by animals //Ecological Modelling. V. 197(3−4). P. 516−519. Elsevier.
- Campagna, C., Bisioli, C., Quintana, F., Perez, F. & Vila, A., 1992. Group breeding in sea lions: pup survive better in colonies. // Animal Behaviour. V. 43. P. 541−548.
- Campagna, C. & Le Boeuf B.J., 1988a. Reproductive behaviour of southern sea lions. // Behaviour. V. 104(3−4). P. 233−261.
- Campagna, C. & Le Boeuf В., 1988b. Thermoregulatory behaviour of southern sea lions and its effect on mating strategies//Behaviour. V.107.P. 72−90. JSTOR.
- Campagna, C., Le Boeuf, B.J. & Cappozzo, H.L., 1988a. Pup abduction and infanticide in southern sea lions. //Behaviour. V. 107(1−2). P. 44−60.
- Campagna, C., Le Boeuf, J. & Cappozzo, H.L., 1988b. Group raids: A mating strategy of male southern sea lions. (Otaria flavescens) // Behaviour. V. 105(3−4). P. 224−248.136
- AnmvxoB A.8. PsnQoovkfiiusHoe noe&deHus chews
- Carey, P., 1992. Agonistic behaviour in female New Zealandfur seals, Arctocephalus forsteri//Ethology. V. 92(1). P. 70−80. John Wiley & Sons.
- Cassini, M., 1999. The evolution of reproductive systems in pinnipeds // Behavioral Ecology. V. 10(5). P. 612. ISBE.
- Clutton-Brock, T., 1989. Review lecture: mammalian mating systems // Proceedings of the Royal Society of London: Series B Biological Ciences. V. 236(1285). P. 339−372. JSTOR.
- Clutton-Brock, T., Green, D., Hiraiwa-Hasegawa, M. &Albon, S.D., 1988. Passing the buck: Resource defence, lekking and mate choice in fallow deer. //Behavioral Ecology and Sociobiology. V. 23. P. 281−296.
- Clutton-Brock, T. & Mcauliffe, K., 2009. Female mate choice in mammals. // Quarterly Review Of Biology. V. 84(1). P. 3−27.
- Clutton-Brock, T., Price, O. & MacColl, A., 1992. Mate retention, harassment, and the evolution of ungulate leks // Behavioral Ecology. V. 3(3). P. 234−242.
- Clutton-Brock, T.H., 1988. Reproductive succes. In: Reproductive succes. Clutton-Brock, T.H. (ed). University of Chicago Press. P. 472−486. (ISBN: 0−226−11 058−3)
- Clutton-Brock, T.H. & Parker, G.A., 1995. Sexual coercion in animal societies. // Animal Behaviour. V. 49. P. 1345−1365.
- Coltman, D.W., Bowen, W.D., Boness, D.J. & Iverson, S.J., 1997. Balance foraging and reproduction in the male harbour seal, an aquatically mating pinniped. (Phoca vitulina) // Animal Behaviour. V. 54(3). P. 663−678.
- Coltman, D. W., Bowen, W.D. & Wright, J.M., 1998. Male mating success in an aquatically mating pinniped, the harbour seal (Phoca vitulina), assessed by microsatellite DNA markers. //Molecular Ecology. V. 7(5). P. 627−638.
- Coltman, D.W., W.D.B. & Wright., J.M., 1999. A multivariate analysis ofphenotype and paternity in male harbor seals, Phoca vitulina, at Sable Island, Nova Scotia. //Behavioral Ecology. V. 10. P. 169−177.
- Costa, D., Croxall, J. & Duck, C., 1989. Foraging energetics of Antarctic fur seals in relation to changes in prey availability. // Ecology. V. 70. P. 596—606.
- Costa, D., Thorson, S., Feldkamp, S., Gentry, R., DeLong, R., Antonelis, G. & Croxall, J., 1985. At sea foraging energetics of three species ofpinnipeds. //Federation Proceedings. V. 44(4). P. 1000.
- Cowlishaw, G. & Dunbar, R.I.M., 1991. Dominance rank and mating success in male primates // Animal Behaviour. V. 41. P. 1045−1056.
- Davies, N., 1991. Mating systems // Behavioural ecology: an evolutionary approach. P. 263−294. Blackwell Scientific.
- Deutsch, C., Haley, M. & Le Boeuf B., 1990. Reproductive effort of male northern elephant seals: estimates from mass loss// Canadian Journal of Zoology. V. 68(12). P. 2580−2593. NRC Research Press.
- Dunbar, R., 1982. Intraspecific variations in mating strategy //Perspectives in ethology. V. 5. P. 385−431. Plenum Press New York.
- Edie A.G. 1977. Distribution and movements ofSteller sea lion cows (Eumetopias jubatus) on a pupping colony. Unpubl. M.S. thesis, Univ. British Columbia, Vancouver. 81 pp.
- Efron, B. & Tibshirani, R.J., 1993. An introduction to the bootstrap. Chapman & Hall/CRC. 436pp. (ISBN: 0−412−4 231−2)
- Emlen, S. & Ornig, L., 1977. Ecology, sexual selection and the evolution of mating systes // Science. V. 197. P. 215−223.
- Farentinos, R., 1971. Some Observation on the Play Behavior of the Steller sea lion // Zeitschrift fur Tierpsychologie. V. 28. P. 428−438.
- Fay, F. H 1982. Ecology and biology of the Pacific walrus, Odobenus rosmarus diver gens llliger//N. Am. Fauna. V. 74. P. 1−279.
- Fay, F. H, Ray, C.G. & A., K.A. 1984, «Time and location of mating and associated behavior in the Pacific walrus, Odobenus rosmarus llliger». NOAA, Technical Report 12, 1984.1. PsnpuuVKrui f^HQS
- Festa-Bianchet, M. & Boag, D., 1982. Territoriality in adult female Columbian ground squirrels // Canadian Journal of Zoology. V. 60(5). P. 1060−1066. NRC Research Press.
- Fieberg, J. & Kochanny, C., 2005. Quantifying home-range overlap: the importance of the utilization distribution // Journal of Wildlife Management. V. 69(4). P. 1346−1359. BioOne.
- Firth, D., 1991. Generalized linear models. In: Statistical Theory and Modeling: In Honour of Sir David Cox. Hinkley, D. V., Reid, N. & Snell, E.J. (ed). London: Chapman and Hall. P. 211−215. (ISBN: 0−412−30 590−9)
- Fiscus, C., 1961. Growth in the Steller sea lion // Journal of Mammalogy.. P. 218−223. JSTOR.
- Ford, R. & Krumme, D., 1979. The analysis of space use patterns // Journal of Theoretical Biology. V. 76(2). P. 125−155. Elsevier.
- Francis, J.M., 1987. Interfemale aggression and spacing in the northern fur seal (Callorhinus ursinus) and the California sea lion (Zalophus californianus). Ph.d. thesis. University of California, Santa Cruz.
- Gentry, R.L., 1973. Thermoregulatory behavior of eared seals. //Behaviour. V. 46(1−2). P. 73−93.
- Gentry, R.L., 1974. The development ofsocial behavior through play in the Steller sea lion. // American Zoologist V. 14(1). P. 391−403.
- Gisiner, R., 1985. Male territorial and reproductive behavior in the Steller sea lion, Eumetopiasjubatus. Ph.d. thesis. University of California, Santa Cruz. 146 pp.
- Greenwood, P., 1980. Mating systems, philopatry and dispersal in birds and mammals. // Animal Behaviour. V. 28. P. 140−162.
- Harestad, A., 1973. Social behaviour in a non-pupping colony ofSteller sea-lion (Eumetopias jubata). M. thesis. University of British Columbia. 92 pp.
- Harestad, A.S. & Fisher, H.D., 1975. Social behavior in a non-pupping colony ofStellersea lions (Eumetopias jubata) // Canadian Journal of Zoology. V. 53. P. 1596−1613.
- Harestad, A., 1977. Seasonal abundance of northern sea lions, Eumetopias jubatus (Schreber), at Mclnnes Island, British Columbia // Syesis. V. 10. P. 173−174.
- Harris, A.S., Young, J.S.F. & Wright, J.M., 1991. DNA fingerprinting of harbour seals (Phoca vitulina concolor), male mating behaviour may not be a reliable indicator of reproductive success. // Canadian Journal of Zoology. V. 69. P. 1862−1866.
- Heath, С. B. 1989. The behavioral ecology of the California sea lion. Ph.d. thesis. University of California, Santa Cruz. California. 275 pp.
- Hoelzel, A.R., LeBoeuf B.J., Reiter, J. & Campagna, С., 1999. Alpha-male paternity in elephant seals. // Behavioral Ecology and Sociobiology. V. 46. P. 298−306.
- Hoffman, J., Boyd, I, Amos, W & Ashley, M., 2003. Male reproductive strategy and the importance of maternal status in the Antarctic fur seal Arctocephalus gazella // Evolution. V. 57(8). P. 1917−1930. Wiley Online Library.
- Hoffman, J. I, Forcada, J., Trathan, P.N. & Amos, W., 2007. Female fur seals show active choice for males that are heterozygous and unrelated. // Nature. V. 445(7130). P. 912−914.
- Hogg, J.T. & Forbes, S.H., 1997. Mating in bighorn sheep: frequent male reproduction via a high-risk «unconventional» tactic. //Behavioral Ecology and Sociobiology. V. 41. P. 33−48.
- Hdglund, J. &Alatalo, R.V., 1995. Leks. Princeton, N.J.: Princeton University Press. 248pp. (ISBN: 0−691−3 727−2)1. Annwxoe A. 8.
- Hood, W.R. & Ono., K.A., 1997. Variation in maternal attendance patterns and pup behavior in a declining population of Steller sea lions (Eumetopias jubatus) // Canadian Journal of Zoology. V. 75(8). P. 1241−1246.
- Humphreys, A.P. & Einon, D.F., 1981. Play as a reinforcer for maze-learning in juvenile rats. // Animal Behaviour. V. 29. P. 259−270.
- Hutchinson, R., Ulrich, R. & Azrin, N., 1965. Effects of age and relatedfactors on the pain-aggression reaction // Journal of Comparative and Physiological Psychology. V. 59(3). P. 365−369. Elsevier.
- Jamieson, G.S. & Fisher., H.D., 1972. The pinniped eye: A review. In: Functional Anatomy of Marine Mammals. Harrison, R.J. (ed). Vol. 1 London: Academic Press. P. 245−261. (ISBN: 0−12−328 001-Х)
- Jennions, M. & Petrie, M., 2000. Why do female mate multiply? A review of the genetic benefits. // Biological Reviews of the Cambridge Philosophical Society. V. 75. P. 21−64.
- Joslyn, W., 1973. Androgen-induce social dominance in infant female rhesus monkeys// Journal of Child Psychology and Psychiatry. V. 14(2). P. 137−145. Wiley Online Library.
- Kahm, M., Hasenbrink, G., Lichtenberg-Frat'e, H, Ludwig, J. & Kschischo, M., 2010. grofit: Fitting Biological Growth Curves with R // Journal of Statistical Software. V. 33(7). P. 1−21.
- Kastelein, R. & Weltz, F., 1990. Distribution, abundance, reproduction and behavior of Steller sea lions (Eumetopias jubatus) in Prince William Sound, Alaska //Aquatic Mammals. V. 15. P. 145−157.
- Keane, В., Creel, S.C. & M., W.P., 1996. No evidence of inbreeding avoidance or inbreeding depression in a social carnivore // Behavioral Ecology. V. 7. P. 480−489.
- Kirkpatrik, M., 1987. The evolutionary forces acting on female mating preferences in polygynous animals. In: Sexual selection: testing the alternatives. Chichester: John Wiley. P. 308.
- Kirkpatrik, M., 1985. The evolution of female choice and male parental investment in polygynous species: the demise of’sexy son'. // American Naturalist. V. 124. P. 309−323.
- Kiyota, M., 2005. Site fidelity, territory acquisition and mating success in male northern fur seals (Callorhinus ursinus). //Mammal Study. V. 30(1). P. 19−27.
- Kiyota, M. & Okamura, H., 2005. Harassment, abduction, and mortality of pups by nonterritorial male northern fur seals. //Journal of Mammalogy. V. 86(6). P. 1227−1236.
- Kroger, R. & Katzir, G., 2008. Comparative anatomy and physiology of vision in aquatic tetrapods // Sensory Evolution on the Threshold: Adaptations in Secondarily Aquatic Vertebrates. P. 121−147.
- La Prade, H. & Graves, H., 1982. Polygyny and female-female aggression in red-winged blackbirds (Agelaius phoeniceus) //The American Naturalist. V. 120(1). P. 135−138. JSTOR.
- Le Boeuf B., 1974. Male-male competition and reproductive success in elephant seals // Integrative and Comparative Biology. V. 14(1). P. 163. Soc Integ Comp Biol.
- Le Boeuf B.J., 1991. Pinniped mating systems on land, ice and in the water: Emphasis on the Phocidae. In: Behaviour of Pinnipeds. Renouf D. (ed). London: Chapman and Hall Ltd. P. 410. (ISBN: 0−412−30 540−2)
- Le Boeuf, B.J. & Briggs, K. T., 1977. The cost of living in a seal harem. //Mammalia. V 41, p. 167−195
- Le Boeuf, B.J. & Peterson, R.S., 1969. Social status and mating activity in elephant seals // Science. V. 163(3862). P. 91−93.
- Lidgard, D., Boness, D., Bowen, W. & McMillan, J., 2005. State-dependent male mating tactics in the grey seal: the importance of body size // Behavioral Ecology. V. 16(3). P. 541. ISBE.
- Lidgard, D.C., Boness, D.J. & Bowen, W.D., 2001. A novel mobile approach to investigating mating tactics in male grey seals (Halichoerus grypus) // Journal of Zoology. V. 255. P. 313 320.
- Ligon, J.D. 1999. The evolution of avian breeding systems. Oxford University Press. Oxford. 504pp. (ISBN: 9 780 198 549 130)
- Loughlin, T., Perlov, A. & Vladimirov, V, 1992. Rangewide survey and estimation of total number of Steller sea lions in 1989. //Marine Mammal Science. V 8. P. 220−239.
- Loy, J., Loy, K, Patterson, D., Keifer, G. & Conaway, C., 1978. The behavior of gonadectomized rhesus monkeys. In: Social Play in Primates. Wilson, E.O. (ed). N.Y.: Academic Press. P. 324. (ISBN: 0−12−652 750−4)
- Lunn, N.J. & Boyd, I.L., 1991. Pupping-site fidelity of Antarctic fur seals at Bird Island, South Georgia. (Arctocephalus gazella) // Journal of Mammalogy. V. 72(1). P. 202−206.1. Алтухов Л.:S
- Mathisen О. A., Baade R. Т., Lopp R.J. 1962. Breeding habits, growth and stomach contents of the Steller sea lion in Alaska // Journal of Mammalogy. V. 43. P. 469−477.
- Maynard Smith, J., 1993. The theory of evolution. Cambridge University Press. 354pp. (ISBN: 0−521−45 128−0)
- McRae, S.B. & Kovacs., K.M., 1989. DNA finderprinting used to determine paternity of hooded seal pups. // Eighth Biennial Conference on the Biology of Marine Mammals. 7−11 December, Asilomar Conference Center, Pacific Grove, CA. P. 42.
- Meier, G.W. & Devanney, V.D., 1974. The ontogeny of play within a society: preliminary analysis. // American Zoologist. V. 14. P. 289−294.
- Merrick, R., 1987. Behavioral and Demographic Characteristics of Northern Sea Lion Rookeries. M. thesis. Oregon State University.
- Merrick, R.L. & Loughlin, T.R., 1997. Foraging behavior ofadult female and young-of-the-year Steller sea lions in Alaskan waters. // Canadian Journal of Zoology. V. 75. P. 776−786.
- Milette, L.L., 1999. Behaviour of Lactating Steller Sea Lons During the Breeding Season: A Comperison between a Declining and Stable Population in Alaska. M. thesis. University of British Columbia.
- Milette, L.L. & Trites, A. W., 2003. Maternal attendance patterns of Steller sea lions (Eumetopias jubatus) from stable and declining populations in Alaska. // Canadian Journal of Zoology. V. 81. P. 340−348.
- Miller, E., 2002. Territorial behavior. In: Encyclopedia of marine mammals. Perrin, W.F., Wu.B. & Thewissen, H. (ed). N.Y.: Academic Press. P. 1235−1243. (ISBN: 0−12−551 340−2)
- Miller, E.H., 1975. Social and evolutionary implications of territoriality in adult male New Zealand fur seals, Arctocephalus forsteri (Lesson, 1828), during the breeding season. // Biology of the Seal. P. 170−187.
- Miller, E.H., 1991. Communication in pinnipeds, with special reference to non-acoustic signalling. In: Behaviour of Pinnipeds. Renouf D. (ed). London: Chapman and Hall Ltd. P. 128−235. (ISBN: 0−12−652 750−4)
- Panksepp, J., 1984. The psychobiology of prosocial behaviors: Separation distress, play and altruism. In: Altruism and Aggression. Social and Biological Origins. Zahn-Waxler, C. (ed). Cambridge University Press. P. 19−57. (ISBN: 0−521−42 367−8)
- Panksepp, J., Siviy, S. & Normansell, L., 1984. The psychobiology of play: Theoretical and methodological perspectives. // Neuroscience & Biobehavioral Reviews. V. 8(4). P. 465−492.
- Perry, E.A. & Amos, W., 1998. Genetic and behavioral evidence that harbor seal (Phoca vitulina) females may mate with multiple males. //Marine Mammal Science. V. 14(1). P. 178 182.
- Peterson, R.S., 1965. Behavior of the northern fur seal. Ph.D. thesis, John Hopkins University, Baltimore, Maryland, 214 pp.
- Peterson, R.S. & Bartholomew, G.A., 1967. The natural history and behavior of the California sea lion. (Zalophus californianus) // American Society of Mammalogists Special Publication. V. 1. P. 79.
- Pike G.C., Maxwell B.E. 1958. The abundance and distribution of the northern sea lion (Eumetopias jubata) on the coast of British Columbia // J. Fish. Res. Bd. Canada. 15(1). P. 5−17.
- Pitcher, K. & Calkins, D., 1981. Reproductive biology of Steller sea lions in the Gulf of Alaska//Journal of Mammalogy. V. 62. P. 599−605. JSTOR.
- Powell, D.A. &Creer, T.L., 1969. Interaction of developmental and environmental variables in shock-elicited aggression. // Journal of Comparative and Physiological Psychology. V. 69. P. 219−225.
- R Development Core Team. 2010. «R: A Language and Environment for Statistical Computing"// Vienna, Austria, url: http. V/www.R-project.ore (ISBN: 3−900 051−07−0)
- Reiter, J., Panken, K.J. & Boeuf, B.J.L., 1981. Female competition and reproductive success in northern elephant seals. //Animal Behaviour. V. 29(3). P. 670−687.
- O.Reynolds, J., 1996. Animal breeding systems // Trends in Ecology & Evolution. V. 11(2). P. 68−72. Elsevier.
- Reynolds, J. & Gross, M., 1990. Costs and benefits of female mate choice: is there a lek paradox? //American Naturalist. V. 136(2). P. 230−243. JSTOR.
- Sandegren, F., 1970. Breeding and maternal behaviour of the Steller sea lion (Eumetopias jubata) in Alaska. M. thesis. University of Alaska, Fairbanks, Alaska.
- Schusterman, R., 1981. Steller sea lion Eumetopias jubatus (Schreber, 1776) // Handbook of marine mammals. V. 1. P. 119−141.
- Schwagmeyer, P., 1988. Scramble-competition polygyny in an asocial mammal: male mobility and mating success // The American Naturalist. V. 131(6). P. 885−892. JSTOR.
- Schwagmeyer, P. & Woontner, S., 1986. Scramble competition polygyny in thirteen-lined ground squirrels: the relative contributions of overt conflict and competitive mate searching // Behavioral Ecology and Sociobiology. V. 19(5). P. 359−364. Springer.
- Sivak, J.G., Howland, H.C., West, J. & Weerheim, J., 1989. The eye of the hooded seal, Cystophora cristata, in air and water. // Journal of Comparative Physiology A: Neuroethology, Sensory, Neural, and Behavioral Physiology. V. 165(6). P. 771−777.
- Sjare, B. & Stirling, I., 1996. The breeding behavior of Atlantic walruses, Odobenusrosmarus rosmarus, in the Canadian High Arctic. // Canadian Journal of Zoology. V. 74(5). P. 897−911.
- Smith, Р.К., 1982. Does play matter? Functional and evolutionary aspects of animal and human play.// Behavioral and Brain Sciences. V. 5. P. 139−184.
- O.Stillman, R.A., Clutton-Brock, Т.Н. & Sutherland, W.J., 1993. Black-holes, mate retention, and the evolution of ungulate leks // Behavioral Ecology. V. 4(1). P. 1−16.
- Stirling, I., 1970. Observations on the behavior of the New Zealandfur seal (Arctocephalus forsteri) // Journal of mammalogy. V. 51. P. 766−778. JSTOR.
- Stirling, I., 1971. Studies on the behaviour on the South Ausralian fur seal, Arctocephalus forsteri (Lesson) I. Annual cycle, postures and calls, and adult males during breeding season // Australian Journal of Zoology. V. 19(3). P. 243−266. CSIRO.
- Stirling, I., 1975. Factors affecting the evolution of social bahaviour in the Pinnipedia // Rapports et Proces-Verbaux des Reunions (Denmark). V. 51(4). P. 766−778. CIEM.
- Stirling, I., 1983. The evolution of mating systems in pinnipeds. In: Recent Advances in the Study of Mammalian Behaviour. Eisenberg, J.F. & Kleinman, D.G. (ed). American Society of Mammalogists, Lawrence, Kansas. P. 489−527.
- Takahashi, L. K & Lore, R.K., 1983. Play fighting and the development of agonistic behavior in male andfemale rats. If Aggressive Behavior. V. 9. P. 217−227.
- Taylor, G., 1980. Fighting in juvenile rats and the ontogeny of agonistic behavior // Journal of Comparative and Physiological Psychology. V. 94(5). P. 953−961. Elsevier.
- Trail, P., 1990. Why shouldlek-breeders be monomorphic?//Evolution. V 44. P. 18 371 852. JSTOR.
- Trivers, R., 1972. Parental investment and sexual selection. Vol. 12 (1). P. 136−179. Aldine.
- Twiss, S.D., 1991. Determinants of male mating success in the grey seal, and comparison of mating systems at two breeding colonies. (Halichoerus grypus). Ph.d. thesis. Department of Zoology, The University of Glasgow, Glasgow.
- Twiss, S.D., Poland, V.F., Graves, J.A., Pomeroy, P.P. 2006. Finding fathers: spatiotemporal analysis of paternity assignment in grey seals (Halichoerus grypus). // Mol. Ecol. V. 15. P. 1939−1953
- Van Parijs, S.M., 2003. Aquatic mating in pinnipeds: A review. // Aquatic Mammals. V. 29(2). P. 214−226.
- Van Parijs, S.M. & Clark, C. W., 2006. Long-term mating tactics in an aquatic-mating pinniped, the bearded seal, Erignathus barbatus. // Animal Behaviour. V. 72(6). P. 12 691 277.
- Vila, B.L., and Cassini, M. H 1990. Aggressiveness between females and mother-pup separation in the southern sea lion, in Chubut, Argentina.//Rev. Chil. Hist. Nat. V. 63 P. 169−176.
- Vladimirov, V.A. (1987), «Age specific reproductive behavior in Northern fur seals on Commander Islands.». NMSF, NOAA, Technical Report 51:113−120, 1987.
- Waterman, J., 2007. Male mating strategies in rodents // Rodent societies: an ecological and evolutionary perspective. P. 27−41. University of Chicago Press Chicago, Illinois.
- Wilkinson, I.S., J., C.S., Duignan, P.J. & Gulland, F.M.D., 2000. Infanticide and cannibalism in the New Zealand sea lion, Phocarctos hookeri. // Marine Mammal Science. V. 16(2). P. 494−500.
- Willson, M.F. & Pianka, E.R., 1963. Sexual selection, sex ratio, and mating system. // American Naturalist. V. 97. P. 405−407.
- Winship, A., Trites, A. & Calkins, D., 2001. Growth in body size of the Steller sea lion (Eumetopias jubatus) // Journal of Mammology. V. 82(2). P. 500−519.
- Wittenberger, J., 1980. Group size and polygamy in social mammals // American Naturalist. V. 115(2). P. 197−222. JSTOR.
- Wolf, J., Kauermann, G. & Trillmich, F., 2005. Males in the shade: habitat use and sexual segregation in the Galapagos sea lion (Zalophus californianus wollebaeki) // Behavioral Ecology and Sociobiology. V. 59(2). P. 293−302. Springer.
- Worton, B., 1989. Kernel methods for estimating the utilization distribution in home-range studies // Ecology. V. 70(1). P. 164−168. Eco Soc America.
- Zar, J., 1974. Biostatistical analysis. Prentice-Hall Englewood Cliffs, NJ. 620pp.