Физико-биогеохимическая модель динамики экосистемы Северного моря
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Диссертация
Апробация работы. Работа проводилась в рамках нескольких европейских проектов и вошла в официальные отчеты. Это следующие проекты: МЕЕСЕ, Marine Ecosystem Evolution in a Changing Environment, EU-FP7, www.meece.euECOOP, European COastal-shelf sea Operational observing and forecasting system, EU-FP6, http://www.ecoop.euRECLAIM, REsolving CLimAtic IMpacts on fish stocks, EU-FP6… Читать ещё >
Содержание
- 1. Задачи изучения Северного моря, данные и подход к построению модели
- 1. 1. Введение
- 1. 2. Общее описание гидродинамики Северного моря
- 1. 3. Биологические процессы в Северном море и их мониторинг
- 1. 4. Подход к построению модели динамики экосистемы Северного моря
- 1. 5. Выводы
- 2. Описание совместной физико-биогеохимической модели ЕСОвМО
- 2. 1. Введение
- 2. 2. Краткое описание физического блока ЕС08М
- 2. 2. 1. Краткое описание блоков НАМЗОМ
- 2. 2. 2. Примеры валидации модели НАМБОМ
- 2. 3. Описание биогеохимического блока ЕСОЭМО
- 2. 4. Постановка задачи моделирования
- 3. Валидация модели ЕСОБМО
- 3. 1. Введение
- 3. 2. Валидация модельного сезонного цикла
- 3. 3. Валидация модели с использованием данных многолетнего мониторинга СРЯ
- 3. 4. Выводы
- 4. Применение ЕС08М0 в изучении динамики экосистемы Северного моря
- 4. 1. Введение
- 4. 2. Изучение сезонной динамики планктона
- 4. 3. Изучение межгодовой динамики планктона
- 4. 4. Пример применения ЕСОБМО для расчета выживаемости личинок икры шпрота
- 4. 5. Выводы
- Заключение
- ПРИЛОЖЕНИЕ А
- Список источников литературы
Список литературы
- Алексеева И.В., Ибраев Р. А. Исследование внутригодовой динамики фитопланктона в Черном море с применением трехмерной физико-биологической модели // Сборник «Мировой океан», ВИНИТИ. 2000. Т. 1, С. 79−86.
- База данных ECOSMO // ECOSMO DATA: Part I and Part II. Материалы проекта RECLAIM. 2007. ftp.uib.no/pub/gfi/corinna/RHCLAIM/read me
- Ибраев P.A. Математическое моделирование термогидродинамических процессов в Каспийском море // Москва: ГЕОС. 2008. ISBN 978−5-89 118−418−3, 128 С.
- Романкевич Е.А., Ветров А. А., Пересыпкин В. И. Органическое вещество Мирового океана// Геология и Геофизика. 2009. Т. 50, № 4, С. 401−411.
- Рябченко В.А., Гальцова В. В., Дворников А. Ю., Нитишинский М. А., Пугалова С. С. Моделирование сезонной изменчивости экосистемы в шельфовой зоне моря Лаптевых // Метеорология и гидрология. 2004. № 6, С. 59−71.
- Тутубалин В. Н., Барабашева Ю. М., Григорян А. А., Девяткова Г. Н., Угер. Е. Г. Математическое моделирование в экологии: Историко-методологический анализ // Москва: «Языки русской культуры». 1999. 208 С.
- Флинт М.В., Суханова И.II. Влияние прибрежных фронтов на структуру и продуктивность пелагических экосистем. В кн. Физические, геологические и биологические исследования океанов и морей // Ред. Шаповалов С. М. М.: «НаучЕ1ый мир» 2010. С. 446−465.
- Alekseeva I., Jarsjo J., Schrum С., Destouni G. Estimation of the Aral sea water budget and sea-groundwater interactions using a coupled sea-ice-groundwater model. ECOSMO model application // Journal of Marine Systems. 2009. V. 76, P. 296−309.
- Anderson T. Plankton functional tvpe modelling-: running before we can walk? // Journalhttp://plankt.oxfordiournals.Org/content/27/l 1/10 ffb08a2b5610560b4e314&kevtvpe2=tf ipsecsha
- Arakawa A., Lamb V.R. Computational design of the basic dynamical processes of the UCLA general circulation model // Methods in Computational Physics. 1977. V. 17, P. 173−265.
- Backhaus J.O. A three-dimensional model for the simulation of shelf sea dynamics // Deutsche Hydrographische Zeitsehrif. 1985. V. 38, P. 165−187.
- Badin G., Williams R. G., Holt J.T., Fernand L. Are mesoscale eddies in shelf seas formed by baroclinic instability of tidal fronts? // Journal of Geophysical Research. 2009. V. 114, № CIO, P. 1−18. doi: 10.1029/2009JC005340.
- Beare D.J., Burns F., Greig A., Jones E.G., Peach K., Kienzle M., McKenzie E., Reid D.G. Long-term increases in prevalence of North Sea fishes having southern biogeographic affinities // Marine Ecology Progress Series. 2004. V. 284, P. 269−278.
- Beaugrand G., Brander K. M., Souissi S. The Influence of Climate Change on North Atlantic Fish Stocks // Poster. ICES symposium, Bergen, Norway. 2004. www.imr.no/2004symposium/index.php
- Beaugrand G., Ibanez F. Monitoring marine plankton ecosystems. II: Long-term changes in North Sea calanoid copepods in relation to hydro-climatic variability // Marine Ecology Progress Series. 2004. V. 284, P. 35−47.
- Beaugrand G., Brander K. M., Lindley J. A., Souissi S., Reid P. C. Plankton effect on cod recruitment in the North Sea // Nature. 2003. V. 426, P. 661−664.
- Beaugrand G., Reid P.C., Ibanez F. Major reorganisation of North Atlantic pelagic ecosystems linked to climate change // GLOBEC International Newsletter. 2002. V. 8, № 2, P. 30−33.
- Benson B.B., Krause D. The concentration and isotopic fractionation of oxygen dissolved in freshwater and seawater in equilibrium with the atmosphere // Limnology and Oceanography. 1984. V. 29, P. 620−632.
- Bergstrom St., Carlssen B. River Runoff to the Baltic Sea: 1950−1990 // Ambio. 1994. V. 23, P. 280−287.
- Brown J., Hill A., Fernand L., Horsburgh K. Observations of a seasonal jet-like circulation at the Central North Sea cold pool margin. Estuarine // Coastal and Shelf Science. 1999. V. 48, P. 343−355.
- Brush M.J., Brawley J.W., Nixon S.W., Kremer J.N. Modeling phytoplankton production: problems with the Eppley curve and an empirical alternative // Marine Ecology Progress Series. 2002. V. 238, P. 31−45.
- Budeus G. Th. A tidal-mixing front in the southern German Bight (in German) // Ph.D. thesis, University of Hamburg. 1989.
- Cloern J.E., Grenz C., Vidergar-Lucas L. An empirical model of the phytoplankton chlorophyll: carbon ratio—the conversion factor between productivity and growth rate // Limnology and Oceanography. 1995. V. 40, № 7, P. 1313−1321.
- Cohen R.E. and Lough R.G. Length-weight relationships for several copepods dominant in the Georges Bank, Gulf of Maine area // Journal of Northwest Atlantic Fishery Science, 1981. V. 2, P. 47−52.
- Corten A., Lindley J.A. The use of CPR data in fisheries research // Progress in Oceanography. 2003. V. 58, P. 285−300.
- Dalrymple R.W. Tidal depositional systems // Facies Models and Sea Level Changes, R. G. Walker and N. P. James, eds.: Geological Association of Canada. 1992. P. 195−218.
- Denman K.L., Pena M.A. The response of two coupled one-dimensional mixed layer/planktonic ecosystem models to climate change in the NE subarctic Pacific Ocean // Deep-Sea Research I. 2002. V. 49, P. 5739−5757.
- Directive 2008/56/EC of the European Parliament and of the Council of 17 June 2008 establishing a framework for community action in the field of marine environmental policy // Marine Strategy Framework Directive. 2008.
- Dommenget D., Latif M. A cautionary note on the interpretation of EOF // Journal of Climate. 2002. V. 15, P. 216−225.
- Eppley R.W. Temperature and Phytoplankton Growth in the Sea // Fishery Bulletin. 1972. V. 70, P. 1063−1085. http://lgmacweb.env.uea.ac.uk/green ocean/publications/Nano/Eppley72.pdf
- Fasham M.J.R., Ducklow H.W., McKelvie S.M.A nitrogen-based model of plankton dynamics in the oceanic mixed layer // Journal of Marine Research. 1990. V. 48, P. 591 639.
- Faure V., Pinazo C., Torreton J.P., Douillet P. Relevance of various formulations of phytoplankton chlorophyll a: carbon ratio in a 3D marine ecosystem model. // Comptes Rendus Biologies. 2006. V. 329, P. 813−822.
- Flynn K. Castles built on sand: dysfunctionality in plankton models and the inadequacy of dialogue between biologists and modellers // Journal of Plankton Research. 2005. V. 27, P. 1207−1210.
- Fofonoff N. P., Millard R. C. Algorithms for computation of fundamental properties of sea water // UNESCO Technical Papers in Marine Science. 1983. 44 PP.
- Franks P.J.S. Phytoplankton blooms at fronts: patterns, scales and physical forcing mechanisms // Reviews in Aquatic Sciences. 1992. Y. 6, P. 121−137.
- Frey H. Stratification during periods of oxygen deficiency in the German Bight during the summers from 1981 to 1983: a comparison with the long-term variation in stratification // Meeresforschung. 1990. V. 32, P. 306−328.
- Fulton E., Smith A., Johnson C. Effect of complexity on marine ecosystem models // Marine Ecology Progress Series. 2003. V. 253, P. 1−16. http://www.int-res.com/abstracts/meps/v253/p1-l 6/
- Gentleman W., Leising A., Frost B., Strom S., Murray J. Functional responses for zooplankton feeding on multiple resources: a review of assumptions and biological dynamics // Deep-Sea Research II. 2003. V. 50, P. 2847−2875.
- Gibson R., Kallberg P., Uppala S. The ECMWF Re-Analysis (ERA) project. // ECMWF Newsletter. 1996. V. 73, P. 7−17.
- Harris G.P. Phytoplankton ecology. Structure, function and fluctuation // London, New York, Chapman and Hall. 1986. 384 PP.
- Harte J. Toward a synthesis of the Newtonian and Darwinian worldviews // Physics Today. 2002. V. 55, 29 PP.
- Hibler W.D. A dynamic thermodynamic sea ice model // Journal of Physical Oceanography. 1979. V. 9, P. 815−846.
- ICES 2009. Report of the Working Group on Modelling of Physical/Biological Interactions (WGPBI)// El Rompido (Iluelva), Spain. ICES CM 2009/0cc:02. 41 PP. http://www.ices.dk/reports/SSGEF/2009/WGPBI09.pdf
- ICES 2005. Guidance on the Application of the Ecosystem Approach to Management of Human Activities in the European Marine Environment // ICES Cooperative Research Report. 2005. № 273, 22 PP.
- ICES 1983. Flushing times of the North Sea.-Coop // ICES Cooperative Research Report. 1983. № 123, 159 PP.
- Janssen F., Schrum C., Backhaus J.O. A climatological data set of temperature and salinity for the North Sea and the Baltic Sea // Deutsche Hydrographische Zeitsehrif. 1999. V. 51, № 9, P. 5−245.
- Janssen F., Schrum C., Hubner U., Backhaus J.O. Validation of a decadal simulation with a regional model for North Sea and Baltic Sea//Climate Research. 2001. V. 18, P. 55−62.
- Johns D.G., Reid P.C. An overview of plankton ecology in the North Sea // Strategic environmental Assessment SEA 2. Technical report TR-005. Saphos. 2006. 29 PP.
- Jungclaus J.H., Botzet M., Haak M., Keenlyside N., Luo J.-J., Latif M., Marotzke J., Mikolajewicz U., Roeckner E. Ocean circulation and tropical variability in the coupled model ECHAM5/MPI-OM // Journal of Climate. 2006. V. 19, P. 3952−3972.
- Kirchman D.L., Rich J.H. Regulation of bacterial growth rates by dissolved organic carbon and temperature in the equatorial Pacific Ocean // Microbial Ecology. 1997. V. 33, P. 11−20.
- Kochergin V. P. Three-dimensional prognostic models. In Three-dimensional Coastal Ocean Models // Coastal and Estuarine Sciences, ed. N. S. Heaps. American Geophysical Union, Washington, D.C. 1987. V. 4, P. 201−208.
- Krause M., Fock H., Greve W., Winkler G. North Sea Zooplankton: A Review // Senkenbergiana Maritima. 2003. V. 33, P. 71−204.
- Krause M., Dippner J.W., Beil J. A review of hydrographic distribution of Zooplankton biomass and species in the North Sea with particular reference to a survey conducted in January-March 1987 // Progress in Oceanography. 1995. V. 35, P. 81−152.
- Krause M., Knickmeyer R. Estimation of the load of cyclic organochlorins in North Sea Zooplankton // Helogolander Meeresuntersuchungen. 1992. V. 46, P. 69−91.
- Krause M., Martens P. Distribution patterns of mesozooplankton biomass in the North Sea // Helgol. wiss. Meeres. 1990. V. 44, P. 295−327.
- Lenhart H.-J., Patsch, J. Daily nutrient loads of the European continental rivers for the years 1977−1998. Berichte aus dem Zentrum fur Meeres- und Klimaforschung- Reihe B // Ozeanographie. 2001. № 40, 146 PP.
- Leppanen A. River data availability in Baltic Sea countries // Internal report FEI. 2002. 8 PP.
- Liebig J. Chemistry in its application to agriculture and physiology // Taylor and Walton, (4th ed., 1947). 1984. 352 PP.
- Liss P. S., Melivat L. Air-sea gas exchange rates: introduction and synthesis. // In P. Buat-Menard, editor, The role of air-sea exchange in Geochemical cycling. 1986. P. 113−127.
- Mesinger F., Arakawa A. Numerical methods used in atmospheric models //JOC, GARP Publication Series. 1976. V. 1, № 17. 64 PP.
- Moll A., Radach G. Review of three-dimensional ecological modelling related to the North Sea shelf system Part 1: Models and their results // Progress in Oceanography. 2003. V. 57, P. 175−217.
- Monod J. Recherches sur la croissance des cultures bacteriennes // Hermann et Cie, Paris. 1942.
- Nielsen T. G., Lokkegaard B., Richardson K., Pedersen F. B., Hansen L. Structure of plankton communities in the Dogger Bank area (North Sea) during a stratified situation // Marine Ecology Progress Series. 1993. V. 95, P. 115−131.
- Nielsen T. G., Richardson K. Food chain structure of the North Sea plankton communities: seasonal variations of the role of the microbial loop // Marine Ecology Progress Series. 1989. V. 56, P. 75−87.
- Neumann T. Towards a 3D-ecosystem model of the Baltic Sea // Journal of Marine Systems. 2000. V. 25, P. 405−419.
- OSPAR Commission 2000. Quality Status Report, Region II Greater North Sea // OSPAR Commission, London. 2000. 136 + xiii PP. http://www.ospar.org/content/content.asp7memF00650830000000 0 0
- Reid P.C., Edwards M., Beaugrand G., Skogen M., Stevens, D. Periodic changes in the zooplankton of the North Sea during the twentieth century linked to oceanic inflow // Fisheries 0ceanography.2003. V. 12, P. 260−269.
- Reid P. C., Borges M. F., Svendsen E. A regime shift in the North Sea circa 1988 linked to changes in the North Sea horse mackerel fishery // Fisheries Research. 2001. V. 50, P. 163−171.
- Radach G., Moll A. Review of three-dimensional ecological modelling related to the North Sea shelf system Part 2: Model validation and data needs // Oceanography and Marine Biology- an Annual Review. 2006. V. 44, P. 1−60.
- Radach G., Patsch J. Climatological annual cycles of nutrients and chlorophyll in the North Sea // Journal of Sea Research. 1997. V. 38, № ¾, P. 231−248. http://www.ifm.7maw.de/rileadmin/files/images/Staff/paetsch/5.pdf
- Richardson A.J., Walne A., Witt M., Lindley J.A., John A. W. G., Sims D. W. Using Continuous Plankton Recorder Data // Progress in Oceanography. 2006. V. 68, P. 27−74.
- Richardson K., Visser A. W., Bo Pedersen F. Subsurface phytoplankton blooms fuel pelagic production in the North Sea // Journal of Plankton Research. 2000. V. 22, P. 16 631 671.
- Ryabchenko V.A., Gorchakov V.A., Fasham M.J.R. Seasonal dynamics and biological productivity in the Arabian Sea euphotic zone as simulated by a three-dimensional ccosystem model // Global Biogeochemical Cycles. 1998. V. 12, № 3, P. 501−530.
- Schrum C., St. John M., Alekseeva I. ECOSMO, a coupled ecosystem model of the North Sea and Baltic Sea: Part II. Spatial-seasonal characteristics in the North Sea as revealed by
- EOF analysis // Journal of Marine Systems. 2006b. V. 61. № 1−2. P. 100−113. doi: 10.1016/i.imarsvs.2006.01.004.
- Schrum C., Siegismund F. 2002. Model configuration of the North Sea Baltic Sea model (in German) // Reports of the Centre for Marine and Climate Research, University of Hamburg, 44 PP.
- Schrum C., Janssen F., Hubner U. Recent climate modelling for North Sea and Baltic Sea. Part A: Model description and validation // Berichte des Zentrums fiir Meeres- und Klimaforschung, Universitat Hamburg. 2000. V. 37, 59 PP.
- Schrum C., Backhaus J.O. Sensitivity of atmosphere-occan heat exchange and heat content in North Sea and Baltic Sea. A comparative Assessment // Tellus 1999. № 51 A. P. 526−549.
- Simpson J.H., Allen C. M., Morris N. C. G. Fronts on the Continental Shelf // Journal of Geophysical Research. 1978. V. 83, P. 4607−4614.
- Skogen M.D., Moll A. Importance of ocean circulation in ecological modeling: An example from the North Sea // Journal of Marine Systems. 2005. V. 57, № 3−4, P. 289 300.
- Skogen M.D., Soiland H., Svendsen E. Effects of changing nutrient loads to the North Sea // Journal of Marine Systems. 2004. V. 46, P. 23−38. doi:10.1016/j.jmarsys.2003.11.013.
- Skogen M.D., Soiland H. Availability of river discharges and river nutrient loads to the North Sea // Internal report IMR. 2001. http://www.imr.no/~morten/nocomments/publications.html
- Skogen M.D., Moll A. Interannual variability of the North Sea primary production: comparison from two model studies // Continental Shelf Research. 2000. V. 20, P. 129 151.
- Sundcrmann J., Beddig S. Zirkulation und Schadstoffumsatz in der Nordsee (ZISCH) // BMBF project MFU 0576 5 (1.1.1988−31.12.1989), Final report University of Hamburg. 1990. 193 PP.
- The Common Fisheries Policy A user’s guide// Luxembourg: Office for Official Publications of the European Communities. 2009. 36 PP. www.ec.europa.eu/fisheries
- Thomas H., Bozec Y., de Baar H. J.W., Elkalay K., Frankignoulle M., Schiettecatte L.-S., Borges A.V. The Carbon budget of the North Sea // Biogeosciences. 2005. V. 2, № 1, P. 87−96.
- Van Aken H.M., van Heijst G.J.F., Maas L.R.M., Observations of fronts in the North Sea // Journal of Marine Research. 1987. V. 45, P. 579- 600.
- Wakelin S. L., Holt J.T., Proctor R. The influence of initial conditions and open boundary conditions on shelf circulation in a 3D ocean-shelf model of the north east Atlantic // Ocean Dynamics. 2009. V. 59, P. 67−81. doi:10.1007/sl 0236−008−0164−3.
- Weston K., Fernand L., Mills D.K. Primary production in the deep chlorophyllaximum of the central North Sea // Journal of Plankton Research. 2005. V. 27, P. 909−922.
- White P.A., Kalff J., Rasmussen J.B., Gasol J.M. The effect of temperature and algal biomass on bacterial production and specific growth rate in freshwater and marine habitats.//Microbial Ecology. 1991. V. 21, P. 99−118.
- Winther N. G., Johannessen J.A. North Sea circulation: Atlantic inflow and its destination //Journal of Geophysical Research. 2006. V. 111, CI 2018, doi: 10.1029/2005JC003310
- Williams R., Lindley J.A. Plankton of the Fladen Ground during FLEX 76 I. Spring development of the plankton community // Marine Biology. 1980. V. 57, P. 73−78.