Разнообразие генетических систем катаболизма нафталина штаммов флуоресцирующих псевдомонад
Выделена новая группа генов nahAc. Разработаны новые специфические праймеры для обнаружения методом ПЦР и характеристики генов nahG и nahR. Впервые изучено разнообразие генов nahG, обнаружены два новых варианта последовательностей гена nahG. Показано, что встречаемость различных вариантов генов nahAc и nahG отличается для штаммов-деструкторов, принадлежащих к разным видам. Обнаружено… Читать ещё >
Содержание
- СПИСОК СОКРАЩЕНИЙ И УСЛОВНЫХ ОБОЗНАЧЕНИЙ
- 1. ОБЗОР ЛИТЕРАТУРЫ
- 1. 1. ОБЩАЯ ХАРАКТЕРИСТИКА ПОЛИЦИКЛИЧЕСКИХ АРОМАТИЧЕСКИХ УГЛЕВОДОРОДОВ
- 1. 2. БИОЛОГИЧЕСКАЯ ДЕСТРУКЦИЯ ПАУ
- 1. 3. БИОХИМИЧЕСКИЕ ПУТИ МЕТАБОЛИЗМА ПАУ БАКТЕРИЯМИ
- 1. 3. 1. Биохимические пути биодеградации нафталина
- 1. 3. 2. Биохимические пути деградации фенантрена
- 1. 4. РАЗНООБРАЗИЕ БАКТЕРИАЛЬНЫХ ГЕНЕТИЧЕСКИХ СИСТЕМ ДЕГРАДАЦИИ ПАУ
- 1. 4. 1. Гены катаболизма ПАУ грам — отрицательных бактерий
- 1. 4. 1. 1. Организация генов катаболизма нафталина плазмиды NAH
- 1. 4. 1. 2. Регуляция генов катаболизма нафталина плазмиды NAH
- 1. 4. 1. 3. Генетические системы катаболизма нафталина аналогичные nah-тенш плазмиды NAH
1.4.1.4. Генетические системы катаболизма нафталина, отличающиеся от архетипа плазмиды NAH7:. а) phd-гены Comamonas testosteroni. б) nag-гены Ralstonia sp. штамм U2. в) /г/ш-гены Burkholderia sp. RP г) гены катаболизма ПАУ бактерий рода Sphingomonas. д) phn-гены Cycloclasticus sp. А5.
1.4.2, Гены катаболизма ПАУ грам — положительных бактерий.
1.4.2.1. war-гены бактерий рода Rhodococcus.
1.4.2.2. Организация генов катаболизма фенантрена Nocardioides sp. КР7.
1.5. УЧАСТИЕ ПЛАЗМИД В БИОДЕГРАДАЦИИ ПАУ.
1.6. ПУТИ ЭВОЛЮЦИИ ГЕНЕТИЧЕСКИХ СИСТЕМ КАТАБОЛИЗМА ПАУ.
2 МАТЕРИАЛЫ И МЕТОДЫ.
2.1. Бактериальные штаммы и плазмиды.
2.2. Питательные среды и условия роста.
2.3. Выделение бактериальных штаммов из почвенных образцов.
2.4 Выделение тотальной ДНКбактерий.
2.5. Выделение плазмидной ДНК.
2.6. Конъюгационный перенос плазмид.
2.7. Полимеразная цепная реакция.
2.8. Гидролиз ДНК эндонуклеазами рестрикции.
2.9. Электрофорез в агарозном геле.
2.10. Препаративное выделение фрагментов ДНК из агарозного геля.
2.11 Мечение ДНК методом рассеянной затравки.
2.12 Гибридизация ДНК на нейлоновых фильтрах.
2.13. .Цитирование ДНК.
2.14. Приготовление компетентных клеток E.coli.
2.15 Трансформация клеток E. coli плазмидной ДНК.
2.16 Определение нуклеотидной последовательности ДНК.
2.17 Определение удельных активностей ферментов.
3. РЕЗУЛЬТАТЫ.
3.1. Изоляция и характеристика штаммов-деструкторов нафталина.
3.2. Генотипический анализ штаммов Р. fluorescens.
3.3. Участие плазмид в генетическом контроле деградации нафталина и фенантрена.
3.4. Анализ удельных активностей ферментов биодеградации нафталина, фенантрена и салицилата.
3.5. Амплификация и RFLP — анализ ключевых генов биодеградации нафталина.
3.6.Анализ генов nahAc штаммов-деструкторов нафталина.
3.7. Анализ генов nahG штаммов-деструкторов.
3.8. Салицилат 5-гидроксилаза в штамме Р. putida АК5.
3.9. Амплификация регуляторного гена nahR.
3.10. Амплификация генов орто- а мета- пути деградации катехола.
3.11. Анализ штаммов на наличие плазмид биодеградации нафталина 1псР-9 группы.
3.12. Геномный фингерпринт (rep-PCR) штаммов-деструкторов нафталина, содержащих Р-9 плазм иды.
3.13. RFLP — анализ плазмид р и 5 -подгрупп Р-9 группы несовместимости.
3.14. Анализ штаммов — деструкторов нафталина на наличие плазмид 1псР-7 группы.
3.15. Локализация генов биодеградации нафталина в штаммах — хозяевах 1псР-7 плазмид.
3.16. RFLP — анализ 1псР-7 плазмид.
4. ОБСУЖДЕНИЕ.
ВЫВОДЫ.
- 1. 4. 1. Гены катаболизма ПАУ грам — отрицательных бактерий
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